The complex enzymatic machinery that assembles cellulose chains spans the plasma membrane, with one part on the cytoplasmic side positioned to bind the substrate, UDP-glucose, and elongate the chains, and another part extending to the outside, responsible for exporting the cellulose molecules to the extracellular space.

Freeze-fracture electron microscopy shows a cellulose synthesis complex, or rosette, composed of six large particles arranged in a regular hexagon with a diameter of about 30 nm. Several proteins, including the catalytic subunit of cellulose synthase, make up this structure. Each of the six particles of the rosette most likely contains three cellulose synthase molecules, each synthesizing a single cellulose chain.

In one working model of cellulose synthesis, cellulose chains are initiated by the transfer of a glucose residue from UDP-glucose to a “primer” glucose already bound to cellulose synthase on the cytoplasmic side of the plasma membrane, to form a disaccharide. As addition of further glucose residues lengthens the chain, it is extruded through a channel formed by the transmembrane helices of cellulose synthase and, on the outer surface of the plasma membrane, joins growing chains from neighboring cellulose synthase molecules to form a cellulose microfibril.

Polymers of more than 6 to 8 glucose units are insoluble in water, promoting microfibril crystallization. There is no definite length for a cellulose polymer; synthesis is highly processive, and some polymers are as long as 15,000 glucose units.

正确

错误

因该企业编制虚假财务会计报告，市财政局由2003年4月5日对其做出罚款3万元的处罚，限15日内缴纳。

光华公司与该企业签订的买卖合同，因企业被宣告破产而终止，光华公司要求赔偿由此造成的损失2万元。

该企业应于2003年6月8日向京普公司支付已购买并入库的材料价款5万元。因企业被人民法院宣告破产，京普公司要求提前支付货款5万元。

该企业厂长以企业名言借用荣和公司一辆轿车供其亲属使用，荣和公司要求返还。

The monosaccharide monomers in cellulose are bonded together differently than in starch.

Cellulose molecules are much too large.

The bonds between the monosaccharide monomers in cellulose are much stronger.

Starch is made of glucose,cellulose is made of fructose.

因该企业编制虚假财务会计报告，市财政局 2003年4月5日对其作出罚款3万元的处罚，限15日内缴纳

光华公司与该企业签订的买卖合同，因企业被宣告破产而终止，光华公司要求赔偿由此造成的损失2万元

该企业应于2003年6月8日向京普公司支付已购买并入库的材料价款5万元，因企业被人民法院宣告破产，京普公司要求提前支付货款5万元

该企业厂长以企业名义借用荣和公司一辆轿车供其亲属使用，荣和公司要求返还

财政局于2004年4月5日对其作出罚款3万元的处罚，限15日内缴纳

光华公司与该企业签订的买卖合同，因该企业被宣告破产而终止，光华公司要求赔偿由此造成的损失2万元

该企业应于2004年6月8日向京海公司支付已购买并入库的材料价款5万元，因企业被人民法院宣告破产，京海公司要求提前支付货款5万元

该企业厂长以企业名义借用荣和公司一辆轿车供其亲属使用，荣和公司要求返还

细胞

蔗糖

纤维素

糖原

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The orientation of the newly deposited extracellular cellulose microfibrils is determined by the orientation of the underlying intracellular microtubules.

The large cellulose synthase enzyme complexes are integral membrane proteins that continuously synthesize cellulose microfibrils on the outer face of the plasma membrane.

The distal ends of the stiff microfibrils become integrated into the texture of the cell wall, and their elongation at the other end pushes the synthase complex along in the plane of the plasma membrane. The microtubules serve as tracks that help guide the movement of the enzyme complexes.

The cortical array of microtubules attached to the plasma membrane by transmembrane proteins helps determine the direction in which the microfibrils are laid down. In this curiously indirect way, the cytoskeleton controls the shape of the plant cell and the modeling of the plant tissues.