The relationship between sentences may be signaled with a sentence connector. Where it is not, the relationship is implicit and you need to work out the connection. What is the relationship between the second and the first sentence? Is it an explanation, contrast, or example? Wetlands can be an important source of protein. The production of fish in shallow ponds or rice paddies developed several thousands of years ago in China and Southeast Asia, and crayfish harvesting is still practiced in the wetlands of Louisiana and the Philippines.